The fossil record of birds is patchy and their evolutionary history is poorly known. The first feathered animal, Archaeopteryx, has been identified in Upper Jurassic deposits, from 150 million years ago (mya). However, while it does appear intermediate between early reptiles and birds, there is some disagreement over whether it is a direct ancestor of present day birds.
Fossils unequivocally of birds do not appear until the Cretaceous period, 80–120 mya, although the number of species suggests that they radiated earlier. The earliest remains are of large flightless diving birds, Hesperornisspp.,with primitive teeth. Other toothed sea birds also lived during the Cretaceous, including the flighted ichthyosaurs.
Also appearing in the Early Cretaceous were the Enantiornithes, a little understood group of seemingly primitive birds. At the end of the period, the toothed birds disappeared with the dinosaurs. Since then, only toothless birds have been found in the record and it is not clear how or when they arose, though it is thought that it was during the Cretaceous.
By the Eocene (c. 50 mya), many modern forms were recognizable. These are non-
Several other forms, mostly large birds, were also present in the Eocene but died out. Other giant birds such as the larger moas of New Zealand and the elephant birds of Africa and Madagascar survived until about 10,000 years ago when they were exterminated by humans.
The evolutionary success of birds is evidenced by the wide variety of present-